1.Department of Ophthalmology, USC Roski Eye Institute, University of Southern California, Los Angeles, CA 91030, USA

2.Departments of Physical Medicine and Rehabilitation, Ophthalmology, Anatomy, and Neurobiology, University of California, Irvine, Irvine, CA 92697, USA

3.Stem Cell Research Center, University of California, Irvine, Irvine, CA 92697, USA

4.USC Ginsburg Institute for Biomedical Therapeutics, University of Southern California, Los Angeles, CA 90033, USA

5.Biomedical Engineering, University of California, Irvine, Irvine, CA 92617, USA

6.Unit on Ocular and Stem Cell Translational Research, National Eye Institute, NIH, Bethesda, MD 20892, USA